Description
Key Characters:
Growth Form:
Perennial, stout, erect or sprawling herbs 5–10(–20?) dm tall, glabrous and glaucous.
Stems:
Caulescent.
Roots:
From a short caudex bearing a circle of several fleshy roots.
Leaves:
Leaves compound (pinnate or ternate-pinnate).
Alternate, distichous, cauline leaves few, like basal ones but reduced upward, the sheaths inconspicuous.
The larger leaves, especially the terminal ones, often 1–3–lobed, 7–13 cm long, 2.5–7 cm wide; ovate to triangular–ovate, 1–3.5 dm long, the leaflets ovate–lanceolate to orbicular.
Apex subacute to obtuse.
Base truncate to obliquely cuneate.
Margins serrate or serrate-dentate.
Palmately veined.
Petiolate to sessile; petioles stout, 1–5 dm long, rather narrowly sheathing at base.
Stipules absent.
Flowers:
Flowers in a fertile terminal umbel and usually 1 or more staminate, lateral, axillary umbels. The terminal peduncle stout, 1–2 dm long, involucre of several linear–oblong to lanceolate bracts up to 2 cm long, these deciduous or sometimes absent. Rays 15–25, spreading-ascending, 5–8 cm long, unequal, webbed, umbellets 10–20–flowered.
Flowers polygamous (with unisexual and bisexual flowers on the same plant). Rays few to numerous, spreading-ascending, often webbed. Mmature pedicels spreading-ascending, 5–12 mm long, unequal, webbed, involucel of 6–8 linear–oblong, scarious–margined bractlets 2–5 mm long.
Calyx lobes relatively prominent, ovate, persistent, ca 2.5 mm long.
Corolla of 5 petals; petals white, orbicular, with a narrower inflexed apex.
Stamens 5, inserted on an epigynous disk.
Ovary inferior, 2-celled; ovules 1 per cells, anatropous; styles 2, usually swollen at base into a stylopodium. Stylopodium conical, exceeding the short styles.
Fruit:
Fruit oblong to oblong–ovoid; 10–13 mm long; 5–8 mm wide; the mericarps flattened dorsally; glabrous; the dorsal ribs low and rounded; the lateral ones broadly winged; the wing narrower to broader than the body; vittae small; 2–4 in the intervals; 5–12 on the commissure; commissural face plane; mericarps readily separating.
Seeds 1 per mericarp; embryo small; endosperm cartilaginous. seed with cotyledons lanceolate; 2.5–4 cm long; 6–12 mm wide; long–petiolate.
Ploidy:
2n = 66*
Habitat:
Occurring on windward cliffs.
Elevation Range:
0–250 m.